Abstract
Full Text
Evolution of Complexity in Paleozoic Ammonoid Sutures
W. B. Saunders, D. M. Work, and S. V. Nikolaeva

Supplementary Material

Appendix Measurements of suture complexity in Paleozoic and Triassic ammonoids, along with stratigraphic occurrences, and phylogenetic ancestors. The following list comprises one set of morphometric data for each of the 598 Devonian-Triassic ammonoid genera included in this study. Chronostratigraphic abbreviations are as follows: 1, Lower Devonian (Emsian); 2, Middle Devonian (Eifelian-Givetian); 3, lower Upper Devonian (Frasnian); 4, upper Upper Devonian (Famennian); 5, lower Lower Carboniferous (Tournaisian-earliest Visean); 6, middle Lower Carboniferous (Visean); 7, upper Lower Carboniferous (lower Namurian); 8, lower Upper Carboniferous (Morrowan); 9, middle Upper Carboniferous (Atokan-Desmoinesian); 10, upper Upper Carboniferous (Missourian-Virgilian); 11, Lower Permian (Asselian-Kungurian); 12, Middle Permian (Roadian-Capitanian); 13, Upper Permian (Wuchiapingian-Changxingian); 14, basal Lower Triassic (lower Griesbachian). SI: sinuosity index; CF: complexity factor; SCI: suture complexity index [= (CF) (SI)]; ANC: ancestor or near ancestor; for further explanation of characters, see (5). Generic and suprageneric classification and phylogenetic relationships generally follow V. E. Ruzhencev, Mollusca-Cephalopoda 1. Superorder Ammonoiddea, Y. A. Orlov, Ed., Fundamentals of Paleontology (Academy of Sciences USSR, Moscow, 1962), vol. 5; ---, M. F. Bogoslovskaya, Trudy, Paleontological Institute, Academy of Sciences USSR (1978), vol. 167; B.I. Bogoslovsky, ibid., vol. 127; B. F. Glenister, W. M. Furnish, in The Ammonoidea: the Evolution, Classification, Mode of Life and Geological Usefulness of a Major Fossil Group, M. R. House, J. R. Senior, Eds. (Academic Press, London, 1981) pp. 49-64; J. Kullmann, ibid., pp. 37-48; M. R. House, ibid. pp. 3-36; E. T. Tozer, ibid., pp. 65-100.
Supplemental Table 1.
TAXONOMY LEVEL SI CF SCI ANC
Order ANARCESTIDA
Suborder AGONIATITINA
Superfamily MIMOSPHINCTACEAE
Family ANETOCERATIDAE
1. Anetoceras 1.17 1.1 1.29
2. Erbenoceras 1.11 1.1 1.22 1
Family MIMOSPHINCTIDAE
3. Luofuceras 1.17 1.1 1.29 1
4. Ruanites 1.17 1.1 1.29 1
5. Mimosphinctes 1.30 1.5 1.95 4
6. Talenticeras 1.25 1.5 1.88 4
Superfamily MIMOCERATACEAE
Family TEICHERTICERATIDAE
7. Lenzites 1.25 1.5 1.88 4
8. Teicherticeras 1.23 1.5 1.85 7
9. Convoluticeras 1.16 1.5 1.74 8
10. Taskanites 1.25 1.5 1.88 7
11. Laganites 1.29 1.5 1.94 9
12. Fasciculoceras 1.27 1.5 1.91 9
Family MIMOCERATIDAE
13. Gyroceratites 1 1.10 1.5 1.65 7
Superfamily AGONIATITACEAE
Family MIMAGONIATITIDAE
14. Mimagoniatites 1.13 1.5 1.70 9
15. Palaeogoniatites 1.07 1.5 1.61 14
16. Parentites 1.29 1.5 1.94 15
17. Kimoceras 1.32 2.0 2.64
Family AGONIATITIDAE
18. Agoniatites 1.47 2.0 2.94 20
19. Paraphyllites 1.27 2.0 2.54 20
20. Chlupacites 1.42 1.5 2.13 26
21. Sellagoniatites 1.50 1.5 2.25 20
Suborder AUGURITINA
Superfamily AUGURITACEAE
Family AUGURITIDAE
22. Gaurites 1.26 2.0 2.52 16
23. Celaeceras 1.64 2.5 4.10 16
Suborder ANARCESTINA
Superfamily ANARCESTACEAE
Family ANARCESTIDAE
24. Anarcestes 1-2 1.25 1.5 1.88 26
25. Atlantoceras 2 1.25 1.5 1.88
26. Archanarcestes 1 1.16 2.0 2.32 14
27. Latanarcestes 1 1.14 2.0 2.28 26
28. Mimanarcestes 1 1.26 1.5 1.89 26
29. Subanarcestes 2 1.27 2.0 2.54 24
30. Paranarcestes 1 1.38 1.5 2.07 24
31. Werneroceras 1-2 1.36 1.5 2.04 33
32. Sellanarcestes 1 1.23 1.5 1.85 26
33. Praewerneroceras 1 1.18 1.5 1.77 27
34. Cabriecoceras 2 1.13 1.1 1.24
35. Archoceras 2-4 1.42 2.0 2.84
Family PINACITIDAE
36. Holzapfeloceras 2 1.18 2.0 2.36
37. Mithraxites 2 1.46 2.0 2.92 39
38. Fidelites 2 1.26 1.5 1.89 20
39. Foordites 2 1.11 1.5 1.67 38
40. Pinacites 2 1.43 2.0 2.86 39
41. Pseudofoordites 2 1.42 2.0 2.84 40
Superfamily PROLOBITACEAE
Family PROLOBITIDAE
42. Sobolewia 2 1.19 1.1 1.31 31
43. Prolobites 2,4 1.25 2.0 2.50 42
44. Renites 4 1.45 2.0 2.90 42
Family PHENACOCERATIDAE
45. Clymenoceras 4 1.74 2.0 3.48 43
46. Cycloclymenia 4 2.02 3.0 6.06 43
Suborder GEPHUROCERATINA
Superfamily GEPHUROCERATACEAE
Family ACANTHOCLYMENIIDAE
47. Ponticeras 3 1.32 2.0 2.64 51
48. Probeloceras 3 1.79 2.0 3.58 50
49. Uchtites 3 1.49 2.0 2.98 47
50. Acanthoclymenia 3 1.49 2.0 2.98 47
51. Pseudoprobeloceras 2-3 1.50 1.5 2.25 35
63. Prochorites 3 1.90 2.0 3.80 50
Family GEPHUROCERATIDAE
52. Sphaeromanticoceras 3 2.28 2.0 4.56 53
53. Manticoceras 3 1.89 2.0 3.78 47
54. Costamanticoceras 3 1.37 2.0 2.74 53
55. Enseites 3 1.37 2.0 2.74 53
56. Trimanticoceras 3 1.89 2.0 3.78 53
57. Delphinites 3 1.60 2.0 3.20 53
58. Playfordites 3 1.89 2.0 3.78 53
59. Mixomanticoceras 3 2.02 2.0 4.02 53
60. Clauseniceras 3 1.72 3.0 3.44 53
61. Maternoceras 3 1.59 2.0 3.18 52
62. Neomanticoceras 3 1.74 2.0 3.48 53
64. Crickites 3 1.42 2.0 2.84 53
65. Gogoceras 3 1.33 2.0 2.66
Family KOENENITIDAE
66. Carinoceras 3 1.63 2.0 3.26 53
67. Koenenites 3 2.09 2.5 5.23 53
68. Hoeninghausia 3 1.82 2.5 4.55 67
69. Timanites 3 1.49 3.0 4.47 68
70. Komioceras 3 1.55 4.5 6.98 53
71. Protimanites 3 1.73 2.5 4.33
Superfamily PHARCICERATACEAE
Family TRIAINOCERATIDAE
72. Tamarites 2 2.18 1.5 3.27
73. Triainoceras 2 2.51 3.5 8.79 72
74. Wellsites 3 2.86 7.5 21.15
75. Sandbergeroceras 2-3 1.90 5.0 9.50 73
76. Schindewolfoceras 2-3 2.26 6.0 13.56 75
Family PHARCICERATIDAE
77. Sphaeropharciceras 2 1.75 2.5 4.38 24
78. Pharciceras 2 2.20 5.0 11.00 77
79. Nordiceras 3 1.45 3.5 5.08 78
80. Synpharciceras 2 2.26 7.5 16.95 82
81. Neopharciceras 2-3 2.49 14.0 34.86 80
82. Stenopharciceras 2 2.03 5.0 10.15 78
Family PETTEROCERATIDAE
83. Petteroceras 2 1.89 7.5 14.18 78
84. Meropharciceras 2 2.58 9.5 24.57
Family DEVONOPRONOITIDAE
85. Devonopronorites 3 2.13 5.7 12.14 78
Superfamily BELOCERATACEAE
Family EOBELOCERATIDAE
86. Eobeloceras 3 1.61 5.0 8.05 61
Family BELOCERATIDAE
87. Mesobeloceras 3 2.20 4.5 9.90 86
88. Beloceras 3 2.54 10.5 26.67 87
Suborder TIMANOCERATACEAE
Family TIMANOCERATIDAE
89. Timanoceras 3 1.38 1.5 2.07 52
Order CLYMENIIDA
Suborder GONIOCLYMENIINA
Superfamily SELLACLYMENIACEAE
Family HEXACLYMENIIDAE
90. Hexaclymenia 4 1.14 1.5 1.71
91. Progonioclymenia 4 1.04 1.5 1.56 90
92. Soliclymenia 4 1.08 1.5 1.62 90
Family COSTACLYMENIIDAE
93. Costaclymenia 4 1.55 2.0 3.10 91
94. Mesoclymenia 4 1.49 2.0 2.99 93
Family SELLACLYMENIIDAE
95. Sellaclymenia 4 1.96 2.0 3.92 93
Family MIROCLYMENIIDAE
96. Uraloclymenia 4 1.09 1.5 1.64 90
97. Pachyclymenia 4 1.16 1.5 1.74 96
98. Borkowia 4 1.11 1.5 1.67
Family BILOCLYMENIIDAE
99. Kiaclymenia 4 1.57 2.0 3.14 96
100. Biloclymenia 4 1.38 2.0 2.76 99
101. Rhiphaeoclymenia 4 1.55 2.0 3.10 99
102. Dimeroclymenia 4 1.30 2.0 2.60
Family WOCKLUMERIIDAE
103. Wocklumeria 4 1.80 2.0 3.60 97
104. Epiwocklumeria 4 1.46 2.5 3.65 103
105. Synwocklumeria 4 1.42 2.5 3.55 104
106. Kielcensia 4 1.43 2.0 2.86
Family GLATZIELLIDAE
107. Glatziella 4 1.14 2.0 2.28 92
108. Postglatziella 4 1.44 2.0 2.88 107
Superfamily GONIOCLYMENIACEAE
Family GONIOCLYMENIIDAE
109. Finiclymenia 4 1.89 3.0 5.67
110. Gonioclymenia 4 2.02 2.5 5.05 94
111. Kalloclymenia 4 2.34 2.5 5.85 110
112. Otoclymenia 4 2.49 2.5 6.23 110
113. Schizoclymenia 4 2.19 3.0 6.57 110
114. Sphenoclymenia 4 1.79 3.5 6.27 110
Superfamily PARAWOCKLUMERACEAE
Family PARAWOCKLUMERIIDAE
115. Kamptoclymenia 4 1.06 2.0 2.12 92
116. Triaclymenia 4 1.13 1.1 1.24 115
117. Parawocklumeria 4 1.19 1.5 1.79 115
Suborder CLYMENIINA
Superfamily CLYMENIACEAE
Family CYRTOCLYMENIIDAE
118. Platyclymenia 4 1.10 1.5 1.65
119. Trigonoclymenia 4 1.10 1.5 1.65
120. Pleuroclymenia 4 1.03 1.5 1.55
121. Spinoclymenia 4 1.07 1.1 1.18 118
122. Flexiclymenia 4 1.13 1.3 1.41
123. Praeflexiclymenia 4 1.05 1.3 1.31
124. Cyrtoclymenia 4 1.06 1.5 1.59 118
125. Stenoclymenia 4 1.06 1.1 1.17 124
126. Trochoclymenia 4 1.18 1.5 1.77 118
127. Piriclymenia 4 1.29 1.1 1.42 118
128. Gyroclymenia 4 1.04 1.1 1.14
129. Protactoclymenia 4 1.10 1.1 1.21
130. Pricella 4 1.18 1.5 1.77
131. Nodosoclymenia 4 1.27 1.1 1.40
Family RECTOCLYMENIIDAE
132. Rectoclymenia 4 1.27 1.1 1.40 118
133. Falciclymenia 4 1.29 1.1 1.42 132
134. Cteroclymenia 4 1.22 2.0 2.44 132
Family CARINOCLYMENIIDAE
135. Carinonoclymenia 4 1.33 1.5 2.00 132
136. Acriclymenia 4 1.96 1.1 2.16 135
137. Pinacoclymenia 4 1.32 2.5 3.30 135
138. Karaclymenia 4 1.17 1.5 1.76 135
Family CLYMENIIDAE
139. Aktuboclymenia 4 1.08 1.1 1.19 118
140. Clymenia 4 1.17 1.1 1.29 141
141. Protoxyclymenia 4 1.63 1.5 2.45 139
142. Kosmoclymenia 4 1.63 1.5 2.45 141
143. Eokosmoclymenia 4 1.11 1.3 1.39
144. Franconiclymenia 4 1.16 1.5 1.74
145. Linguaclymenia 4 1.33 1.5 1.99
146. Muessenbiaergia 4 1.40 2.0 2.80
Family CYMACLYMENIIDAE
147. Genuclymenia 4 1.29 2.0 2.58 139
148. Kazakhoclymenia 4 1.49 2.5 3.73 150
149. Liroclymenia 4 1.9* 1.1* 2.10*
150. Cymaclymenia 4 1.44 1.5 2.16 147
151. Ornatoclymenia 4 1.22 1.5 1.83 150
152. Laganoclymenia 4 1.31 1.5 1.97 150
153. Siekluckia 4 1.24 1.5 1.86
Order GONIATITIDA
Suborder TORNOCERATINA
Family MAENIOCERATIDAE
154. Maenioceras 2 1.55 2.5 3.88 24
155. Afromaenioceras 2 1.55 2.5 3.88 24
Family TORNOCERATIDAE
156. Protornoceras 4 1.20 2.0 2.40 158
157. Epitornoceras 3 1.20 2.0 2.40 158
158. Tornoceras 2-4 1.26 2.0 2.52
159. Aulatornoceras 2-4 1.24 2.0 2.48 158
160. Tornia 4 1.03 1.5 1.58 156
161. Armatites 4 1.34 2.0 2.68 168
162. Pseudoclymenia 4 1.51 2.0 3.02 173
163. Kirsoceras 4 1.15 1.5 1.73 168
164. Wedekindella 2 1.28 2.0 2.56
165. Lobotornoceras 3 1.27 3.0 3.80 158
166. Domanikoceras 3 1.26 2.0 2.52 156
167. Crassotornoceras 3-4 1.29 2.0 2.58 158
168. Polonoceras 4 1.28 2.0 2.56 159
169. Gundolficeras 4 1.56 2.0 3.12 175
170. Exotornoceras 4 1.95 2.5 4.88 169
171. Linguatornoceras 3-4 1.56 2.0 3.12 158
172. Simicheiloceras 4 1.35 2.0 2.70 156
173. Pernoceras 4 1.16 2.0 2.32 168
174. Trevoneites 2 1.09 2.0 2.18
175. Falcitornoceras 4 1.18 2.0 2.36 176
176. Phoenixites 2-4 1.21 2.0 2.42 158
177. Truyolsoceras 4 1.49 2.0 2.98 159
178. Oxytornoceras 4 1.55 2.0 3.10 156
183. Parodiceras 2 1.49 2.0 2.98 29
Family SINOTITIDAE
179. Sinotites 4 1.28 2.0 2.56
Family POSTTORNOCERATIDAE
180. Posttornoceras 4 2.13 3.0 6.39 170
181. Discoclymenia 4 2.06 3.2 6.59 180
182. Alpinites 4 2.18 4.0 8.72 180
Family CHEILOCERATIDAE
184. Melonites 4 1.10 2.0 2.20 186
185. Raymondiceras 4 1.10 2.0 2.20 186
186. Cheiloceras 4 1.28 2.0 2.56
187. Dyscheiloceras 4 1.72 2.0 3.44 186
188. Torleyoceras 4 1.27 2.0 2.54 186
189. Paratorleyoceras 4 1.53 2.0 3.06 188
Family SPORADOCERATIDAE
190. Sporadoceras 4 1.77 3.0 5.31 188
191. Maeneceras 4 1.89 3.0 5.67 190
192. Araneites 4 1.68 3.0 5.04 190
193. Nothosporadoceras 4 1.59 3.0 4.77 190
Family DIMEROCERATIDAE
194. Paradimeroceras 4 1.71 2.5 4.28 189
195. Dimeroceras 4 1.59 2.0 3.18 194
196. Paratornoceras 4 1.67 2.0 3.34 195
197. Praemeroceras 4 1.53 2.0 3.06
198. Acrimeroceras 4 1.85 2.0 3.70
Superfamily PRAEGLYPHIOCERATACEAE
Family PRAEGLYPHIOCERATIDAE
199. Praeglyphioceras 4 1.98 2.5 4.95 200
200. Lagowites 4 1.68 2.0 3.36 188
Superfamily PRIONOCERATACEAE
Family PRIONOCERATIDAE
201. Prionoceras 4 1.41 2.0 2.82
202. Balvia 4 1.41 2.0 2.82 209
203. Acutimitoceras 5 1.61 2.0 3.22 204
204. Stockumites 4-5 1.78 2.0 3.56 209
205. Costimitoceras 5 1.54 2.0 3.08 208
206. Cunitoceras 6 1.80 2.0 3.60 207
207. Imitoceras 5 1.82 2.0 3.64 208
208. Rectimitoceras 4-5 1.66 2.0 3.32
209. Mimimitoceras 4-5 1.91 2.0 3.82 201
210. Nicimitoceras 5 1.62 2.0 3.24 208
211. Paragattendorfia 5 1.77 2.0 3.54 204
212. Globimitoceras 5 1.81 2.0 3.62 209
213. Irinoceras 5-7 1.83 2.0 3.66 207
214. Gattendorfia 5 1.64 2.0 3.28 204
215. Gattenpleura 5 2.08 2.0 4.16 214
216. Kazakhstania 5 2.08 2.0 4.16 214
217. Zadelsdorfia 5 1.75 2.0 3.50 214
218. Pseudarietites 5 1.72 2.0 3.44 219
219. Paprothites 5 1.60 2.0 3.20
220. Paralytoceras 5 1.66 2.0 3.32 218
Family MAXIMITIDAE
221. Maximites 9-10 1.56 2.0 3.12
Family PSEUDOHALORITIDAE
222. Neoaganides 10-13 1.49 2.0 2.98 221
223. Pseudohalorites 11 2.00 2.6 5.20 222
224. Zhonglupuceras 11 1.58 2.8 4.42 229
225. Shouchanogoceras 12 1.53 2.0 3.06 231
226. Aulacaganides 11 1.94 2.0 3.88 222
227. Elephantoceras 12 1.60 2.0 3.20
228. Erinoceras 12 1.57 2.0 3.14
229. Lianyuanoceras 11 1.71 2.0 3.42 222
230. Qinglongites 13 1.31 2.0 2.62 222
231. Sanghzhites 12 1.62 2.0 3.24
232. Shangraoceras 12 1.56 2.0 3.12 229
233. Sosioceras 12 1.65 2.0 3.30 222
234. Yinoceras 11 2.82 4.0 11.28 223
235. Lanceoloboceras 12 2.31 5.0 11.55 223
Family KARAGANDOCERATIDAE
236. Karagandoceras 5 2.24 2.5 5.60 210
237. Voehringerites 5 1.61 2.0 3.22 203
Family QIANNANITIDAE
238. Qiannanites 5 1.97 3.0 5.91 220
239. Asidoceras 5 2.15 5.5 11.83 238
Family PRODROMITIDAE
240. Eoprodromites 5 1.82 3.3 6.01 238
241. Prodromites 5 1.96 12.1 23.72 240
Suborder GONIATITINA
Superfamily PERICYCLACEAE
Family PERICYCLIDAE
242. Goniocyclus 5 1.63 2.0 3.26
243. Pericyclus 5 1.82 2.0 3.64 242
244. Rotopericyclus 5 1.82 2.0 3.64 243
245. Asiacyclus 5 1.85 2.0 3.70 243
246. Neopericyclus 5 1.87 2.0 3.74 244
247. Polaricyclus 5 1.59 2.0 3.18 243
248. Isopericyclus 5 1.56 2.0 3.12
249. Rhiphaeocyclus 6 1.95 2.0 3.90
250. Fascipericyclus 5 2.27 2.0 4.54 243
251. Fascipericyclus-2 5 2.27 2.0 4.54 243
252. Hammatocyclus 5-6 2.20 2.0 4.40
253. Helicocyclus 5 2.13 2.0 4.16
254. Ammonellipsites 5 2.61 2.5 6.53 250
255. Stenocyclus 5 1.74 2.0 3.48 247
Family MUENSTEROCERATIDAE
256. Muensteroceras 5 1.65 2.0 3.30 242
257. Bollandites 6 1.80 2.0 3.60 252
258. Bollandoceras 6 2.20 2.0 4.40 260
259. Cluthoceras 7 1.41 2.0 2.82 261
260. Dzhaprakoceras 5-6 1.81 2.0 3.62 256
261. Beyrichoceratoides 5-7 1.96 2.0 3.92 256
262. Eurites 5 2.30 2.0 4.60 256
263. Terektytes 5 1.96 2.0 3.92 256
Family INTOCERATIDAE
264. Intoceras 5 1.27 2.0 2.54 265
265. Aquilonites 5 1.58 2.0 3.16 256
266. Oxintoceras 6 1.27 2.0 2.54 267
267. Quasintoceras 5-6 1.18 2.0 2.36 264
Family KOZHIMITIDAE
268. Kozhimites 5 1.64 2.0 3.28
Family FURNISHOCERATIDAE
269. Furnishoceras 5 1.90 2.0 3.80
Superfamily DIMORPHOCERATACEAE
Family DIMORPHOCERATIDAE
270. Dimorphoceras 6 2.04 2.9 5.92
271. Asturoceras 7 2.51 2.8 7.03 272
272. Trizonoceras 6-7 2.22 2.4 5.34 270
273. Sulcodimorphoceras 7 2.11 2.9 6.12
274. Paradimorphoceras 6-7 2.53 3.3 8.39 270
275. Glyphiolobus 6-8 1.82 2.9 5.28 270
276. Metadimorphoceras 6-8 2.95 3.3 9.73 270
277. Anthracoceratites 8 2.25 3.2 7.20
Family BERKHOCERATIDAE
278. Kazakhoceras 6-7 2.11 2.7 5.69
Family ANTHRACOCERATIDAE
279. Anthracoceras 7 1.44 2.0 2.88 280
280. Beyrichoceras 6 1.83 2.0 3.66
281. Sudeticeras 6-7 1.81 2.5 4.53 280
Family EOGONIOLOBOCERATIDAE
282. Winchelloceras 5-6 1.93 2.0 3.86
283. Eogonioloboceras 6-7 1.68 2.5 4.20 282
284. Arcanoceras 6-7 1.83 2.5 4.58 283
Family GIRTYOCERATIDAE
285. Girtyoceras 6 1.94 2.5 4.85 280
286. Eumorphoceras 7 2.40 2.5 6.0 288
287. Peytonoceras 7 1.43 2.0 2.86 286
288. Sulcogirtyoceras 6 1.81 2.5 4.53 285
289. Tumulites 7 1.72 2.0 3.44 286
290. Torulites 6 1.67 2.0 3.34
291. Cowdaleoceras 6 1.96 2.0 3.92
Family UNKNOWN
292. Zhifangoceras 5 2.06 2.5 5.15
Superfamily NOMISMOCERATACEAE
Family NOMISMOCERATIDAE
293. Nomismoceras 6 1.58 2.0 3.16 295
294. Eonomismoceras 5 1.79 2.0 3.58 253
295. Simmonoceras 5 1.61 2.0 3.22 294
296. Beleutoceras 7 1.48 2.0 2.96 293
297. Cavilentia 7 1.53 2.0 3.06 293
298. Hudsonoceras 7-8 1.58 2.5 3.95 293
299. Baschkirites 8 1.59 2.5 3.98 298
Family ENTOGONITIDAE
300. Entogonites 6 1.79 2.0 3.58 293
Superfamily GONIATITACEAE
Family GONIATITIDAE
301. Goniatites 6 1.97 2.5 4.93 257
302. Arnsbergites 6 1.91 2.5 4.78 301
303. Goniatitella 6 1.95 2.0 3.90 301
304. Hibernicoceras 6 1.86 2.5 4.65 301
305. Hypergoniatites 6 2.19 2.5 5.48 301
306. Lusitanoceras 6 1.74 2.5 4.35 301
307. Neogoniatites 6-7 1.97 2.5 4.93 304
308. Paraglyphioceras 6 2.03 2.5 5.08 301
Family DELEPINOCERATIDAE
309. Delepinoceras 7 2.17 2.8 6.08 310
310. Platygoniatites 6-7 1.82 2.6 6.50 301
Family AGATHICERATIDAE
311. Dombarites 6-7 2.26 2.5 5.65 301
312. Paracravenoceras 7 2.53 2.5 6.33 302
313. Pericleites 7 2.07 2.8 5.80 311
314. Proshumardites 7-9 2.69 4.5 12.11 311
315. Agathiceras 9-12 2.45 4.5 11.03 314
Family SYGAMBRITIDAE
316. Sygambrites 6 1.95 2.5 4.88
Superfamily NEOGLYPHIOCERATACEAE
Family NEOGLYPHIOCERATIDAE
317. Neoglyphioceras 6-7 1.73 2.0 3.46 327
318. Lusitanites 6-7 1.59 2.0 3.18 317
319. Lytheoceras 7 1.73 2.0 3.46 317
320. Mirilentia 7 1.49 2.0 2.98 318
Family CRAVENOCERATIDAE
321. Cravenoceras 7 1.91 2.0 3.82 327
322. Emstites 6-7 1.69 2.0 3.38 327
323. Alaoceras 6-7 1.88 2.0 3.76 327
324. Caenolyroceras 7 1.69 2.0 3.38 327
325. Dombarigloria 6 2.19 2.0 4.38 327
326. Lyrogoniatites 6-7 1.62 2.0 3.24 327
327. Pachylyroceras 6 1.78 2.0 3.56
328. Quasicravenoceras 7 1.57 2.0 3.14 326
329. Tympanoceras 7 1.82 2.0 3.64 327
330. Collectoceras 7 1.80 2.0 3.60 321
331. Verancoceras 7 1.44 2.0 2.88 328
332. Kardailites 7 1.68 2.5 4.20
333. Nuculoceras 7 1.44 2.0 2.88
334. Aenigmatoceras 8 1.61 2.5 4.03
Family RHYMMOCERATIDAE
335. Rhymmoceras 7 1.65 2.5 4.13 337
336. Fayettevillea 7 1.79 2.5 4.48 335
337. Ophilyroceras 6 1.91 2.0 3.82
338. Chumazites 8 1.66 2.5 4.15
Family FERGANOCERATIDAE
339. Ferganoceras 6 1.63 2.0 3.26
340. Nummoceras 6 1.80 2.5 4.50 339
Superfamily SOMOHOLITACEAE
Family GLAPHYRITIDAE
341. Glaphyrites 7-11 1.98 2.5 4.95 342
342. Richardsonites 7 1.62 2.5 4.05 321
343. Pseudoglaphyrites 7 1.96 2.5 4.90 342
344. Syngastrioceras 7-9 2.27 2.5 5.68 341
Family STENOGLAPHYRITIDAE
345. Stenoglaphyrites 7-8 1.52 2.5 3.80
346. Rhadinites 7 1.73 2.5 4.33 342
347. Euroceras 7 1.75 2.5 4.38 346
348. Physematites 7 1.56 2.5 3.90 346
349. Subitoceras 7 1.69 2.5 4.23 346
350. Zephyroceras 7 1.79 2.5 4.48 346
351. Schartymites 8 1.77 2.5 4.43 346
352. Paraschartymites 8 1.62 2.5 4.05 351
Family PSEUDOPARALEGOCERATIDAE
353. Pseudoparalegoceras 9 2.39 2.5 5.98 354
354. Phaneroceras 8-9 2.49 2.5 6.22
Family CLISTOCERATIDAE
355. Clistoceras 9 1.72 2.5 4.30 345
356. Neogastrioceras 8-9 2.16 2.5 5.40
Family DUNBARITIDAE
357. Dunbarites 9-10 1.66 2.6 4.32 353
Family SOMOHOLITIDAE
358. Somoholites 7-11 2.54 2.5 6.35 344
359. Neoshumardites 11 2.08 2.8 5.82 360
360. Andrianovia 11 2.00 2.8 5.60 358
Family UNCERTAIN
361. Neoglaphyrites 10-11 2.00 2.5 5.00 341
Superfamily NEODIMORPHOCERATACEAE
Family RAMOSITIDAE
362. Cravenoceratoides 7 1.44 2.0 2.88
363. Ramosites 7-8 1.66 2.5 4.15 362
Family NEODIMORPHOCERATIDAE
364. Cymoceras 8 2.16 2.5 5.40 363
365. Schuichengoceras 8 1.91 2.7 5.16 364
366. Dimorphoceratoides 8 1.98 2.7 5.35 365
367. Politoceras 9 1.88 2.7 5.08 365
368. Neodimorphoceras 9-10 2.00 2.7 5.40 367
Superfamily GASTRIOCERATACEAE
Family HOMOCERATIDAE
369. Homoceras 8 1.84 2.5 4.60 371
370. Parahomoceras 8 1.62 2.5 4.05 371
371. Isohomoceras 7-8 1.86 2.5 4.65 342
372. Isohomoceras 7-8 1.79 2.5 4.47 342
373. Vallites 7-8 1.83 2.5 4.58 371
374. Bashkortoceras 8 1.64 2.5 4.10 369
375. Umbetoceras 8 1.51 2.5 3.78 369
376. Fallacites 8 1.59 2.5 3.98 369
377. Bogdanoceras 8 1.86 2.5 4.65 369
Family DECORITIDAE
378. Decorites 8 1.94 2.5 4.85 369
379. Kushanites 8 1.42 2.5 3.55 369
380. Brevikites 8 1.55 2.0 3.10 369
Family SURENITIDAE
381. Surenites 8 1.68 2.5 4.20 369
382. Verneuilites 8 1.94 2.5 4.85 381
383. Marianoceras 8 2.11 2.5 5.27 382
384. Aphantites 8 1.41 2.5 3.52 381
385. Melvilloceras 8 1.85 2.5 4.62 383
386. Gaitherites 8 2.14 2.5 5.35 382
Family RETICULOCERATIDAE
387. Phillipsoceras 8 1.54 2.5 3.85 381
388. Alurites 8 1.57 2.5 3.93 387
389. Tectiretites 8 1.69 2.5 4.23 387
390. Retites 8 1.76 2.5 4.40 387
391. Arkanites 8 1.88 2.5 4.70 390
392. Quinnites 8 2.05 2.5 5.12 390
393. Reticuloceras 8 1.63 2.5 4.07 387
394. Bilinguites 8 1.97 2.5 4.93 393
Family GASTRIOCERATIDAE
395. Monitoceras 8 1.51 2.5 3.78 381
396. Cancelloceras 8 2.29 2.5 5.72
397. Agastrioceras 8 1.96 2.5 4.90 396
398. Gastrioceras 8-9 1.99 2.5 4.97 396
399. Trettinoceras 9 2.22 2.5 5.55
400. Owenoceras 9 1.80 2.5 4.50
Superfamily THALASSOCERATACEAE
Family BISATOCERATIDAE
401. Homoceratoides 8 1.80 2.5 4.50 369
402. Bisatoceras 8-10 1.80 2.5 4.50 401
Family THALASSOCERATIDAE
403. Eothalassoceras 9-10 2.01 3.6 7.24 402
404. Prothalassoceras 10-11 2.05 4.3 8.82 403
405. Gleboceras 10 1.77 3.6 6.37
406. Aristoceras 10-11 1.82 3.6 6.55 405
407. Thalassoceras 11-12 3.71 6.0 22.26 404
408. Epithalassoceras 12 4.83 4.1 19.80 407
409. Artistoceratoides 12 2.25 4.1 9.23 406
Superfamily SCHISTOCERATACEAE
Family SCHISTOCERATIDAE
410. Branneroceras 8 2.42 2.5 6.05 396
411. Diabloceras 8-9 2.46 2.5 6.15 410
412. Paralegoceras 9 2.49 3.0 7.47 411
413. Eoschistoceras 9-10 2.99 5.5 16.44 412
414. Schistoceras 10 2.88 3.6 10.37 413
415. Paraphaneroceras 8 2.28 2.5 5.70 411
416. Inzeroceras 8 2.62 3.0 7.86 410
Family WELLERITIDAE
417. Winslowoceras 9 2.62 3.5 9.17
418. Eowellerites 9 2.66 3.6 9.58 417
419. Wellerites 9 3.14 7.0 21.98 418
Family CHRISTIOCERATIDAE
420. Christioceras 9 3.05 5.1 15.55
Family AXINOLOBIDAE
421. Axinolobus 8 2.89 3.2 9.25 415
Family ORULGANITIDAE
422. Yakutoceras 8 2.09 2.5 5.22
423. Kayutoceras 8 2.13 2.5 5.33 422
424. Parayakutoceras 8 1.92 2.5 4.80 423
425. Yakutoglaphyrites 8 2.23 2.5 5.57 424
426. Aldanites 8 2.11 2.5 5.27
427. Orulganites 8 2.10 2.5 5.25 426
Superfamily GONIOLOBOCERATACEAE
Family GONIOLOBOCERATIDAE
428. Gonioloboceratoides 9 1.69 2.5 4.23 435
429. Gonioloboceras 10 1.98 2.5 4.95 428
430. Mescalites 11 2.15 2.7 5.81 429
Family GONIOGLYPHIOCERATIDAE
431. Gonioglyphioceras 9 1.57 2.5 3.93 428
432. Mangeroceras 9 1.73 2.5 4.32 433
433. Donetzoceras 8-9 1.61 2.5 4.03
434. Wewokites 9 1.57 2.5 3.93
Family WIEDEYOCERATIDAE
435. Wiedeyoceras 8-9 1.70 2.5 4.25
Family UNCERTAIN
436. Pennoceras 10 1.54 2.5 3.85
Superfamily NEOICOCERATACEAE
Family NEOICOCERATIDAE
437. Neoicoceras 9 2.52 2.5 6.30
438. Trochilioceras 9-10 2.10 2.5 5.25 437
439. Eupleuroceras 10 1.38 2.0 2.76 438
440. Eoasianites 10-11 2.41 2.5 6.03 437
Family METALEGOCERATIDAE
441. Metalegoceras 11-12 2.39 3.5 8.37 443
442. Bransonoceras 11-12 2.72 3.5 9.52 441
443. Juresanites 11 2.37 3.0 7.11 440
444. Pseudoschistoceras 11-12 2.54 4.0 10.16 441
445. Clinolobus 11-12 1.72 2.5 4.30
446. Eothinites 11 2.66 3.5 9.31 452
447. Epiglyphioceras 12 1.88 2.5 4.70 452
448. Spirolegoceras 12 2.23 3.5 7.81 449
449. Sverdrupites 12 2.54 3.5 8.89 451
Family PARAGASTRIOCERATIDAE
450. Paragastrioceras 11 2.24 2.5 5.60 452
451. Epijuresanites 11 2.30 2.6 5.98 455
452. Svetlanoceras 11 1.90 2.5 4.75 440
453. Synuraloceras 11 2.30 2.5 5.75 452
454. Tumaroceras 11 2.11 2.5 5.28 450
455. Uraloceras 11 2.17 2.5 5.43 450
456. Atsabites 11 2.19 2.5 5.48 464
457. Anatsabites 12 1.56 2.5 3.90 456
458. Pseudogastrioceras 13 1.77 2.5 4.43 459
459. Altudoceras 12-13 2.01 2.5 5.03 450
460. Aulacogastrioceras 12 2.08 3.0 6.24 464
461. Chekiangoceras 12 2.20 2.5 5.50 450
462. Daubichites 12 2.10 2.5 5.25 464
463. Roadoceras 12-13 2.01 2.5 5.03 464
464. Stenolobulites 11-12 1.75 2.5 4.38 452
465. Strigogoniatites 12-13 2.08 2.5 5.20 459
Superfamily SHUMARDITACEAE
Family SHUMARDITIDAE
466. Preshumardites 10 2.20 2.8 6.16 341
467. Pseudaktubites 10 2.41 3.0 7.23 466
468. Shumardites 10 2.85 5.4 15.39 467
Family PARASHUMARDITIDAE
469. Aktubites 9 1.92 2.8 5.38 341
470. Parashumardites 10 2.61 5.5 14.35 469
471. Eoshumardites 10 2.51 4.7 11.80 469
472. Eovidrioceras 10 2.17 2.8 6.08 469
Family PERRINITIDAE
473. Perrinites 11-12 4.36 13.4 58.42 474
474. Metaperrinites 11 4.29 11.0 47.29 475
475. Properrinites 11 4.20 9.8 41.16 470
Superfamily MARATHONITACEAE
Family MARATHONITIDAE
476. Kargalites 10-11 2.32 6.7 15.54 478
477. Subkargalites 9-10 2.64 6.4 16.90 341
478. Cardiella 10-11 2.36 6.9 16.28 477
479. Marathonites 10 2.75 7.8 21.45 477
480. Almites 11 2.05 6.6 13.53 479
481. Cardiella 10-11 2.33 6.4 14.91 477
482. Pseudovidrioceras 12 2.44 4.8 11.71 480
483. Suakites 11 3.07 6.3 19.34 480
Family HYATTOCERATIDAE
484. Hyattoceras 12-13 4.49 9.9 44.45 485
485. Eohyattoceras 11-12 3.42 7.3 24.97 483
Superfamily CYCLOLOBACEAE
Family VIDRIOCERATIDAE
486. Vidrioceras 10 2.05 5.9 12.10 472
487. Glassoceras 12 3.29 10.4 34.22 488
488. Martoceras 11-13 2.57 8.4 21.59 486
489. Peritrochia 12 2.38 5.8 13.80
490. Prostacheoceras 11-12 2.49 5.7 14.19 486
491. Stacheoceras 11-13 2.69 11.1 29.86 488
492. Tabantalites 11 2.21 6.1 13.48 486
Family CYCLOLOBIDAE
493. Cyclolobus 13 8.49 38.0 323 497
494. Changhsingoceras 13 4.65 14.3 66.50
495. Demarezites 12 5.27 10.1 53.23 503
496. Timorites 12-13 8.39 24.2 203 497
497. Waagenoceras 12 6.24 16.8 105 495
498. Shengoceras 12 4.57 10.2 46.61
499. Guiyangoceras 12 2.78 6.9 19.18 490
500. Liuzhouceras 12 3.66 7.0 25.62
501. Mexicoceras 12 4.54 11.6 52.66 503
502. Paratongluceras 12 4.19 7.1 29.75
503. Tongluceras 12 4.34 11.1 48.17
Superfamily POPANOCERATACEAE
Family POPANOCERATIDAE
504. Popanoceras 11-12 2.77 9.6 26.59 506
505. Epitauroceras 13 2.92 10.3 30.08 508
506. Propopanoceras 11 2.27 7.7 17.48 507
507. Protopopanoceras 11 1.79 6.1 10.92
508. Tauroceras 12 2.67 12.3 32.84 504
Family MONGOLOCERATIDAE
509. Mongoloceras 12 3.19 10.3 32.86
Superfamily ADRIANITACEAE
Family ADRIANITIDAE
510. Emilites 10-11 1.86 3.1 5.77 355
511. Adrianites 12 2.91 7.0 20.37 517
512. Aricoceras 11-12 3.07 5.0 15.35 513
513. Crimites 11-12 2.44 5.5 13.42 510
514. Doryceras 12 2.42 4.5 10.89 519
515. Epadrianites 12-13 3.55 7.5 26.65 517
516. Neoaricoceras 12 3.06 8.2 25.09 513
517. Neocrimites 11-12 2.05 5.6 11.48 513
518. Palermites 12 3.35 6.5 21.78 519
519. Pamiritella 11 2.58 7.0 18.06 513
520. Pseudagathiceras 12 2.93 6.5 19.05 517
521. Pseudoemilites 11 2.15 3.1 6.67 510
522. Sizilites 12 2.74 5.0 13.70 519
523. Sosiocrimites 11-12 3.02 7.5 22.65 517
524. Veruzhites 11 2.21 5.0 11.05 521
525. Hoffmannia 12 3.09 6.0 18.54
526. Texoceras 12 1.82 4.5 8.19
527. Nevadoceras 11 2.44 4.5 10.98 521
Order PROLECANITIDA
Suborder PROLECANITINA
Superfamily PROLECANITACEAE
Family PROLECANITIDAE
528. Eocanites 5 2.46 2.0 4.92
529. Protocanites 5 2.44 2.5 6.10 528
530. Acrocanites 5 2.86 4.8 13.73 529
531. Merocanites 5-6 2.82 3.5 9.87 529
532. Cantabricanites 6 3.32 4.0 13.28 531
533. Michiganites 5-6 2.64 3.5 9.24 529
534. Prolecanites 6-7 2.31 4.0 9.24 533
535. Dombarocanites 6-7 2.35 5.0 11.75 534
Family DARAELITIDAE
536. Epicanites 6-7 3.11 6.1 18.97 533
537. Boesites 8-11 2.41 5.5 13.26 536
538. Daraelites 11-12 2.44 6.9 16.84 537
Superfamily MEDLICOTTIACEAE
Family PRONORITIDAE
539. Pronorites 6 2.33 5.2 12.12 533
540. Megapronorites 6-7 3.09 6.4 19.78 539
541. Metapronorites 9-11 2.74 7.7 21.10 543
542. Pseudopronorites 8-10 3.76 7.7 28.95 543
543. Stenopronorites 7-9 2.88 5.2 14.98 539
544. Uralopronorites 7 2.54 6.7 17.02 539
545. Neopronorites 10-11 3.07 8.2 25.17 541
546. Parapronorites 11-12 3.37 7.7 25.95 541
547. Sakmarites 11 2.29 5.0 11.45 545
Family MEDLICOTTIIDAE
548. Uddenites 10 3.31 10.4 34.42 551
549. Daixites 10-11 3.10 10.3 31.93 551
550. Neouddenites 11 2.86 13.9 39.75 548
551. Prouddenites 10 3.17 9.0 28.53 544
552. Uddenoceras 10 2.98 9.2 27.42 548
553. Medlicottia 11-12 4.39 14.5 63.65 554
554. Artinskia 10-11 4.45 14.7 65.41 552
555. Eumedlicottia 11-13 4.51 19.3 87.04 553
556. Neogeoceras 12-13 4.88 15.6 76.19 553
557. Propinacoceras 11-12 3.69 11.6 42.80 552
558. Akmilleria 11 3.64 14.0 50.96 552
559. Bamyaniceras 11-12 4.25 17.9 76.07 557
560. Difuntites 12-13 2.46 7.0 17.22 557
561. Miklukhoceras 11 4.20 11.2 47.04
562. Sicanites 11-12 4.40 14.5 63.80 565
563. Artioceras 11 4.56 13.1 59.74 552
564. Artioceratoides 11 3.64 13.7 49.87 563
565. Synartinskia 11-12 4.18 15.7 65.63 552
566. Episageceras 13-14 4.41 13.1 57.72
Family SUNDAITIDAE
567. Sundaites 13 3.66 13.2 48.31 546
Order CERATITIDA
Superfamily XENODISCACEAE
Family PARACELTITIDAE
568. Paraceltites 12-13 1.64 3.0 4.92 538
569. Cibolites 12-13 1.75 3.9 6.82 568
570. Nielsenites 12 1.64 5.4 8.86 568
571. Doulingoceras 13 1.60 2.5 4.00 568
Family XENODISCIDAE
572. Xenodiscus 12-14 1.79 4.3 7.70 569
573. Tapashanites 13 2.56 4.8 12.29
574. Tompophiceras 14 1.96 4.6 9.02
Family PSEUDOTIROLITIDAE
575. Pseudotirolites 13 2.25 4.8 10.80
576. Pleuronodoceras 13 2.24 4.1 9.18
577. Rotodiscoceras 13 2.95 5.0 14.75
578. Pentagonoceras 13 2.75 5.2 14.30
Family DZHULFITIDAE
579. Paratirolites 13 2.65 4.5 11.93
Superfamily OTOCERATACEAE
Family ANDERSSONOCERATIDAE
580. Anderssonoceras 13 1.56 2.0 3.12
581. Pericarinoceras 13 1.79 2.5 4.48
582. Lenticoceltites 13 1.79 2.0 3.58
Family ARAXOCERATIDAE
583. Kingoceras 12-13 1.65 3.6 5.94 569
584. Eoaraxoceras 13 3.80 1.9 7.03 569
585. Anfuceras 13 1.98 3.6 7.13
586. Prototoceras 13 1.93 5.2 10.04 584
587. Araxoceras 13 2.06 4.9 10.09 584
588. Dzhulfoceras 13 2.38 10.7 25.47
589. Pseudotoceras 13 2.82 4.7 13.25
Family OTOCERATIDAE
590. Otoceras 14 3.33 7.6 25.31
591. Anotoceras 14 3.33 7.6 25.31
Superfamily NORITACEAE
Family OPHICERATIDAE
592. Ophiceras 14 2.09 5.9 12.33
593. Vishnuites 14 1.64 4.3 7.05
594. Wordieoceras 14 2.18 7.0 15.26
595. Discophiceras 14 2.38 7.8 18.56
596. Hypophiceras 14 1.98 4.7 9.31
Family PROPTYCHITIDAE
597. Bukkenites 14 3.33 5.5 18.32
598. Proptychites 14 1.97 5.2 10.09