COVER A bloom of Rafflesia tuan-mudae from western Borneo. Rafflesiaceae species produce the world's largest flowers, with blooms up to 1 meter in diameter. Molecular evidence described on page 1812 shows that these enigmatic parasites are members of the spurge family, Euphorbiaceae, hence the enormous flowers of Rafflesiaceae most likely arose from tiny-flowered ancestors. Photo: © Ch'ien Lee

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