COVER Rodhocetus, a whale that lived 47 million years ago, visualized on the basis of new Eocene fossils from Pakistan. The ankle bones indicate a close relationship of early whales to hooved land mammals such as hippopotami and pigs. Forefeet retain hooves on the central digits, but hind feet with slender webbed toes indicate that Rodhocetus was predominantly aquatic. [Painting by John Klausmeyer, University of Michigan Exhibit Museum]

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