COVER The enzyme RNA polymerase II in the act of transcribing a gene. X-ray crystal structure comprises the protein (gray, except for orange "clamp" and green "bridge" helix), DNA (blue template strand, green nontemplate strand), and RNA (red). The pink sphere is an active center Mg²⁺ ion. Double-stranded DNA enters from the right and unwinds before the active center. The unwound nontemplate DNA strand is obscured by motion or disorder. 1876 [Adapted from Fig. 2C of Gnatt et al.]

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